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Mak 70th Grad PhD Citations Session 1, Part 1



College of Agricultural and Environmental Sciences (CAES)


PARIYO Anthony (RIP)

Resolution of genetic structure for resistance to cassava brown streak disease: germplasm diversity, resistance stability and inheritance patterns.

Mr. PARIYO Anthony (RIP) studied the genetic structure for resistance to cassava brown streak disease (CBSD), an acute disease that has for decades limited optimal cassava productivity in eastern and southern Africa, which is incited by cassava brown streak viruses (CBSVs). Lack of information on genetic diversity, stability and the mode of inheritance of resistance to CBSD severely limits efforts tailored towards its control. The study resulted into three principal conclusions to guide future work: 1) low frequency of CBSD resistance alleles in eastern Africa cassava germplasm; 2) different CBSD resistance gene actions for foliar and root plant parts; 3) significant environments effects, with Namulonge located in central region, being the most suitable location for CBSD resistance screening. This study was funded by the world Bank coded Millennium Science Initiative, through Government of Uganda, and was supervised by Prof. Phinehas Tukamuhabwa and Dr. Yona Baguma.


ACHORA Janet Cox

Use of Information and Communication Technologies in conservation agriculture knowledge pathways among smallholder farmers in Machakos and Laikipia counties, Kenya

Ms. ACHORA Janet Cox examined the use of Information Communication Technologies (ICTs) in conservation agriculture knowledge pathways among smallholder farmers in Machakos and Laikipia counties, Kenya. To suggest ways in which emerging ICTs can be integrated for enhanced conservation agriculture knowledge sharing, the study identified that the fragmented conservation agriculture knowledge network, the minor role of ICT actors in conservation agriculture knowledge sharing, the low integration of ICT tools and ICT capacities of conservation agriculture actors, constrained the optimal use of ICTs in conservation agriculture knowledge sharing. The study established that an integrated ICT knowledge sharing framework could improve and integrate the use of emerging ICTs for conservation agriculture knowledge sharing. This study was self-funded and partly supported by the Regional Universities Forum for Capacity Building in Agriculture (RUFORUM), and was supervised by Dr. Haroon Sseguya and Dr. Florence Birungi Kyazze.



Modelling the impact of stunting on childhood survival in Buhweju District and the cost of its prevention

Mr. BUKUSUBA John studied the risk factors for the high rate of stunting in Buhweju District, and modelled the impact of its reduction on child survival and the cost of interventions required. The study found half of the children under 5 years were stunted and boys were more stunted than girls. Stunting was largely attributed to low coverage of child survival interventions, morbidity, low consumption of animal-source foods, food insecurity, poverty, and lack of knowledge about stunting. The cost for the reduction of stunting was estimated at US$ 21.2 million for the period 2018-2030, necessitating an additional US$ 250,000 per year for scale up. The interventions and costed strategy can be adapted for the 5-year district development plan and the development of a new multi-sectoral nutrition policy and national development plan. This study was funded by Nestlé Foundation, and was supervised by Prof. Archileo N. Kaaya and Dr. Abel Atukwase.


BYAKIKA Stellah (Ms)

Studies on the safety of Obushera and probiotic potential of selected lactic acid bacteria

Ms. BYAKIKA Stellah examined the safety of Obushera, a popular fermented cereal-based beverage from Uganda. This was motivated by the increasing uncontrolled commercial production of Obushera which compromises consumer safety. She also evaluated the potential contribution of three lactic acid bacteria isolated from Obushera towards improving product safety and human health. Findings showed presence of virulent, antibiotic-resistant and acid-tolerant Escherichia coli and aflatoxins in some Obushera sold in Kampala. The isolates; Lactobacillus plantarum MNC 21, Lactococcus lactis MNC 24 and Weisella confusa MNC 20 bound aflatoxins, inhibited Escherichia coli and exhibited the potential to reduce blood cholesterol, stimulate insulin release and stabilize heart pressure. Adoption of the isolates in processing of Obushera and related products could improve safety and health. The study was co-funded by Mr. Samuel K. Byakika and the Food Technology and Business Incubation Center, and was supervised by Assoc. Prof. Ivan Muzira Mukisa and Prof. Charles Muyanja.


GEBREMEDHN Hailay Mehari

Genetic Resistance to Soybean Rust (Phakopsora Pachyrhizi) in Line UG-5

Mr. GEBREMEDHN Hailay Mehari studied the genetic resistance to soybean rust in line UG-5. Soybean rust (SBR) is a devastative foliar diseases causing high yield losses worldwide. In Uganda, a local line UG-5 seems to have unique genes showing potential contribution towards improvement of SBR, but genetic control of its resistance is not yet characterized. The study revealed significant GCA effects and high Baker’s ratio, suggesting the predominance of additive gene action in the inheritance of SBR resistance. Three putative QTLs were identified on chromosomes 6, 9 and 18. The QTL detected on chromosome 9 was novel and has not been reported elsewhere. Plant defense signaling pathway-related candidate genes were predicted from the QTLs on chromosomes 9 and 18, which could facilitate efficient MAS and gene pyramiding for the development of durable resistance to SBR. This study was funded by INTRA-ACP Mobility Scheme, and was supervised by Prof. Phinehas Tukamuhabwa and Dr. Tomas L. Odong.



Resistance spectrum to anthracnose disease and the genetic effect of pyramided genes on yield in common bean

Mr. KIRYOWA Moses determined the pathogenic variability of Colletotrichum lindemuthianum (fungus causing anthracnose disease); assessed the effectiveness of pyramided resistance genes against anthracnose disease; and determined genetic effect of pyramided genes on yield in beans. C. lindemuthianum was highly variable with 24 new physiological races, worthy of attention. Pyramided genes in the right combination conferred broad-spectrum resistance but with a yield penalty. Some single genes conferred broad-spectrum resistance but may not be durable. The decision to pyramid resistance genes should, therefore be weighed against the threat the pathogen poses. The effective single and pyramided genes are a resource for breeders and pathologists. This study was funded by the National Agricultural Research Organization (NARO) and the International Centre for Tropical Agriculture (CIAT) -Uganda and was supervised by Prof. Phinehas Tukamuhabwa and Dr. Stanley Nkalubo.


KUMI Frank

Studies of sorghum resistance to downy mildew (Peronosclerospora sorghi) disease in Uganda

Mr. KUMI Frank studied the prevalence, distribution and population structure of downy mildew disease in major sorghum growing districts in Uganda. He also characterized Peronosclerospora sorghi isolates which causes downy mildew disease. His study found Arua, Namutumba and Pallisa as downy mildew disease hotspots. In addition, temperature, relative humidity and rainfall were the main drivers for downy mildew epidemics. He further screened Uganda sorghum germplasm for sources of resistance to downy mildew disease under different disease pressure. Two parents and seven crosses were identified as good transmitters of resistance to downy mildew disease and are recommended as lines in a sorghum breeding programme targeting downy mildew resistance and yield improvement. This study was funded by Intra-ACP CSAA project and RUFORUM, and supervised by Prof. Patrick Rubaihayo and Dr. Moses Biruma.



Genetics of cowpea resistance to bruchid (Callosobruchus maculatus Fab.)

Mr. MIESHO Belay studied Bruchids (Callosobruchus maculatus), which is one of the most destructive insect-pests of cowpea causing significant losses in storage. The study was designed to contribute to the reduction of cowpea storage losses through elucidation of cowpea genetics of resistance to bruchids. Through intensive phenotyping, seed biochemical and inheritance studies, four cowpea genotypes (2419, WC42, TVu-2027and IT84s-2246) resistant to bruchids were identified and recommended as donor parents for cowpea breeding against bruchids. Furthermore, eleven genomic regions and six candidate genes associated with the resistance traits were identified using genome-wide association study which could be used for marker assisted breeding. This study was funded by DAAD and Carnegie through RUFORUM; and University of California through MaRCCI; and was supervised by Prof. Patrick Rubaihayo and Prof. Samuel Kyamanyawa.


MSISKA Mercy Ulemu

Genetic resistance to adzuki bean bruchid in soybean

Ms. MSISKA Mercy Ulemu studied the genetic resistance to adzuki bean bruchid (Callosobruchus chinensis) in soybean. Utilization of resistant varieties to manage bruchids is obstructed by lack of sources of resistance and information on genetics of inheritance. Msiska’s study established sources, basis and inheritance of resistance to C. chinensis in soybean. Two genotypes; AVRDC G8527 and PI G89 were identified as sources of resistance. High tannins, total antioxidants, peroxidase activity and low flavonoids were biochemicals associated with resistance. Potential parents in breeding for resistance were SREB-15C, S-Line 9.2 and S-Line 13.2A. Crosses of the SREB-15C x S-Line 13.2A and SREB-15C x Maksoy 3N were recommended as start up material for the breeding programme. This study was funded by Intra ACP-CSAA, APPSA and Carnegie Corporation of New York through RUFORUM, and was supervised by Prof. Phinehas Tukamuhabwa and Prof. Samuel Kyamanywa.



Common bean (Phaseolus vulgaris, L.) extrusion cooking: process optimization and product evaluation

Ms. NATABIRWA Hedwig researched on extrusion of biofortified common beans, with focus on improving the nutritional quality of expanded starchy snack foods, commonly consumed by children. Her work showed that extruded bean product properties were associated with the bean chemical components, and influenced by the extrusion cooking conditions used. She optimized extrusion conditions, thus producing a bean snack with high protein and iron content, and improved protein digestibility. Her research proved that biofortified common beans can be used to produce highly acceptable and nutritive puffed snacks, thus promoting increased intake of macro- and micro-nutrients. The developed process provides an alternative and economically feasible extrusion processing methodology that can be used by industry for production of nutritious expanded snack products.  The study was funded by the ADB-GOU HEST research corporation through CIAT and supervised by Prof. John H Muyonga and Assoc. Prof. Dorothy Nakimbugwe.



Adaptation and genetic analysis of earliness and yield component traits of yam bean (Pachyrhizus Spp.) in Rwanda

Mr. NDIRIGWE Jean studied the adaptation and genetic analysis of earliness and yield component traits of yam bean (Pachyrhizus Spp.) in Rwanda. Yam bean, a high yielding and rich root crop in Latin America and Asia was recently introduced into Rwanda for integration into the diverse farming agro-ecologies and improvement of diets of root crop dependent communities. Ndirigwe’study identified high genetic variability, heritability and significant high general ability, specific ability and their variance components indicating that both additive and non-additive genes control earliness traits and expected genetic gain could be expected in genetic improvement of yam beans introduced. High yielding and well adapted genotypes were AC 209033, AC 209035 and EC209018 and were recommended for participatory farmer’s selection. This study was funded by the Belgium Technical Cooperation, and was supervised by Prof. Phinehas Tukamuhabwa and Prof. Emeritus Patrick Rubaihayo.


ONYILO Francis

Functional genetics in ascomycetes Pseudocercospora fijiensis (Synonym Mycosphaerella fijiensis) the pathogen of black sigatoka disease in banana.

Mr. ONYILO Francis studied Functional Genetics in ascomycetes Pseudocercospora fijiensis (formerly Mycosphaerella fijiensis) the pathogen of black Sigatoka in Banana.  Francis isolated and characterised Mitogen activated protein kinase encoding genes; PfHog1, PfSlt2 and PfFus3 which are responsible for growth and virulence   of Pseudocercospora fijiensis. He developed an RNA interference mediated gene silencing mechanism and an Agrobacterium tumefaciens mediated transformation procedure to introduce genes into mycelium fragments of Pseudocercospora fijiensis. His research is a step towards developing banana cultivars with resistance to black Sigatoka disease. This will help increase overall banana production. Functional genetic tools developed by Francis can be used to investigate other fungal pathogens example Magnaporthe oryzae of Rice. This Study was funded by Norman Borlaug Leadership Enhancement in Agricultural Programme University of California Davis USA and Agricultural Biotechnology Support programme II – USAID. His research was conducted under the mentorship of Prof. Bryce Falk and Prof. Ioannis Stergiopoulos. He was supervised by Dr. Tusiime Geoffrey and Dr. Leena Tripathi.


OPOLOT Henry Nakelet

Unraveling critical factors for a responsive university-farming community engagement in Uganda: insights from two outreach projects at the School of Agricultural Sciences, Makerere University

Mr. OPOLOT Henry Nakelet explored factors for development of a systematic long-term engagement framework between universities and farming communities. Findings showed that: the quality of teaching using participatory methods for development of lifelong skills; field attachment duration and appropriateness of host organizations to support practical learning; timely sharing of information; and farmers’ capacity development are critical for enhancing the role of universities on agricultural development. The study recommended strengthening application of participatory teaching methods for students to develop lifelong learning skills, increasing field attachment duration in accredited organizations for enhanced practical learning, integration of ICTs for timely information sharing, and mainstreaming farmer training into outreach activities. The study also suggested introduction of a post-graduation apprenticeship as an avenue for professional skills development and long-term engagement. This study was funded by RUFORUM CARP03 Project, and was supervised by Dr. Prossy Isubikalu & Dr. Bernard Obaa.



Urban resilience to climate extremes in Mbale municipality in Eastern Uganda

Mr. Oriangi George investigated historic and projected occurrence of precipitation extremes up to the year 2050, assessed factors perceived to be influential in enhancing resilience and proposed and tested a Municipality Resilience Index (MRI) to measure household resilience to precipitation extremes in Mbale municipality. Findings revealed that extreme precipitation periods have become and are likely to become more frequent between September and January. Additionally, household ability to meet its daily expenditure needs, household size, networks with NGOs, health care, education, access to credit and employment showed to be crucial factors in enhancing resilience to precipitation extremes. The MRI revealed that Mbale municipality has a low resilience index (0.2). Thus, contributing to previous studies on community resilience and bear relevance for practitioners to understand where to invest more resources to enhance resilience. This study was funded by SIDA and was supervised by Assoc. Prof. Yazidhi Bamutaze, Dr. Paul Isolo Mukwaya and Prof. Petter Pilesjö.



Comparative analysis of genotypic diversity among Xanthomonas campestris pv. musacearum and Xanthomonas vasicola pv. vasculorum strains

Mr. WASUKIRA Arthur conducted genome wide association studies on bacterial wilt isolates from banana, ensete and sugarcane from Eastern Africa. Bacterial wilt continues to reduce banana productivity and therefore affect livelihoods. He determined two major clades of bacterial strains within Eastern Africa, sequenced fourteen Xanthomonas strains and identified thirty-two specific candidate genes. The genome assemblies are useful in molecular dating studies, specific genetic markers used in functional, epidemiological and biogeographical research. New breeding technologies use pathogen-derived effectors as molecular probes to identify resistance genes for genetic plant protection. This study contributes to transgenic development of banana bacterial wilt resistance through gene editing. The study was funded by Millennium Science Initiative/NARO, University of Exeter, and was supervised by Dr. Geoffrey Tusiime and Dr. Jerome Kubiriba.


Please click the links below to navigate to the PhD Citations for the respective Sessions.

< Director’s Message | Session 1: Part1: Part2 : Part3 | Session 2 | Session 3 | Session 4 >


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Government Sponsorship Admission Lists 2022/23 Verified by Districts



Students hold a group discussion in the Arts Quadrangle, College of Humanities and Social Sciences, Makerere University, Kampala Uganda. Date taken: 13th April 2018.

The Office of the Academic Registrar Makerere University is pleased to announce that the following  candidates have been verified by their respective Districts for admission to the  programmes indicated against their names under the District Quota Scheme.

Follow the link below for the list:

Batch I

Students verified by Districts for the Programmes 2022/23 Academic Year

Batch II

Students verified by Districts for the Programmes 2022/23 Academic Year

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Student Registration for Semester I 2022/2023



(a) First Years
Every new student admitted to a programme of study of Makerere University was issued a provisional admission letter with fees structure for payment of requisite fees. This enables privately sponsored first year students pay at least 60% tuition and all functional fees before issuance of original admission letters which should be collected from the respective Colleges/Schools.

For a candidate to qualify to be a bonafide student of the University, he/she MUST be
registered. Registration is a mandatory requirement of the University which must be
done within the specified time at the beginning of the semester. Failure to do so will
automatically lead to your place being forfeited to another candidate. Official
Registration/Verification of documents is on going using the Academic Information
Management System (ACMIS)
used by Makerere University.

Ensure that you complete all the required registration formalities within the prescribed
time as per the Fees Payment Policy and registration programmes provided by your
respective Colleges. The system cycle will be closed on 3Qth November, 2022.

Registration Requirements
For registration purposes all first year students MUST produce their Original documents
as indicated on their admission letters for validation and verification purposes. At the end
of the online registration exercise, new students will be required to submit 3 photocopies
of their academic documents which will be dully signed and stamped by their Registrars
for record purposes.

(b) Continuing Students
Continuing students also use the Academic Information Management System (ACMIS) for
registration for Academic Year 2022/2023. Continuing students should register online by
accessing the registration Menu in the Student Portal and selecting the first option labeled
“Self Registration” and click the REGISTRATION NOW option.

The Cycle for online registration for the Academic year 2022/2023, Semester One is open
for all continuing students. The system cycle will be closed on 30th November, 2023.

(c) Students who belong to the under listed categories are advised to contact their College/School Registrars before they can register.

(i) Retakes Cases
(ii) Stay Put Cases
(iii) Withdrawal cases
(iv) Audited Courses
(v) Extension Cases
N. B. Each student should pay National Council for Higher Education fee of 20,000/ = per year and UNSA Subscription of 2,000/= per year before registration.

(d) In case of any problem, consult your College/School Registrar. College Accountants are responsible for providing the financial status to all students and generating lists of paid up students to the Deans. They will also clear paid up students to be issued the examination permits before sitting University examinations for Semester One, 2022/2023 Academic year.

Prof. Buyinza Mukadasi

Download the Communication from Academic Registrar here

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A Delegation from Netherlands Visits Makerere University



On Monday 14th November 2022, a delegation from the Royal Kingdom of the Netherlands visited Makerere University to discuss capacity building, scholarships, research and approaches to developing the Technical and vocational education and training (TVET) and Higher Education programme.

The visiting delegation consisted of Mr Siemen Tuinstra, Deputy Director, Department of Social Development; Mr Theodore Klouvasa, Coordination Policy Officer, Education & Youth Responsible for the development of the new TVET & Higher Education Programme; Ms Hilde de Bruijn – Senior Policy Officer and Ms Joy Acom-Okello, the Policy Officer Humanitarian Affairs and Migration at the Netherlands Embassy in Kampala.

Discussion with the Vice Chancellor

The Vice Chancellor, Prof. Barnabas Nawangwe welcomed the visitors to Makerere University and briefed them about the history of the University that started as a technical college in 1922 with 14 students. In 1949, it became a University College affiliated to the University College of London, offering courses leading to the general degrees of its then mother institution. With the establishment of the University of East Africa in June 29, 1963, the special relationship with the University of London came to a close and degrees of the University of East Africa were instituted. On July 1, 1970, Makerere became an independent national university of the Republic of Uganda, offering undergraduate and postgraduate courses leading to its own awards. In 1990, there was liberalization of university education after the World Bank and IMF decided that there should be less spending on university education and introduced structural adjustment programmes. The Government pays a lump sum to the university to sponsor some students and the rest are private students.

The Coordination Policy Officer, Mr Theodore Klouvasa informed the Vice Chancellor about the new programme on TVET and Higher Education that their government was developing. The purpose of their visit was to consult other stakeholders in higher education such as universities, ministries of Education and Sports, Agriculture, Gender and Youth and technical institutions to learn more about the existing collaborations between them and see where the Netherlands government can assist in developing a beneficial programme. How exactly do universities relate with Vocational Institutes and what is the education system in Uganda ad how do donors communicate with the major actors in the education system? How do universities relate with the private sector? If government sponsors some students, how can the scholarships be more inclusive and target the marginalized? Research is very important for all universities. How can they bring more research in the university and what can they add on the PhD infrastructure? Makerere University is strategic partner with the Netherlands having trained many PhDs at Wageningen University, Maastricht University, Vrije Universiteit Amsterdam; University of Groningen; Radboud University Nijmegen; Delft University of Technology.

Makerere University has many collaborations globally and has over the years increased partnerships with the government. The College of Health Sciences has done extensive research with the Military in the area of HIV/AIDS; with the Ministry of Water & Mineral Development in the area of water qualities and management; with UNRA with joint research and use of technologies for materials and road construction; with Ministry of Agriculture and Animal Industry and Fisheries in the area of cross boundary animal diseases; with Food processing industries with our School of Food Nutrition and Biotechnology; the Horticulture industries in controlling quality of products for export; the IT companies with our College of Computing and Information Sciences and also the College of Engineering, Design, Art and Technology. The University relates well with the Uganda Society of Architects and our architecture students are exposed to the new products on the market such as the new design of roofing tiles. The students share simple technologies learnt in class with the manufacturing companies which have helped in boosting production.

Uganda is affected by a high population growth and many graduates cannot find jobs. The education system needs to be geared towards problem solving techniques to be taught to learners/students at all levels. There is a need to change the mindset of the teachers/professors and the students as well. A mindset programme is to be introduced in the first year of studies for all programmes. Makerere University is also in the process of establishing an incubation hub where the good ideas of students can be developed to start a business. If you want to change the country, you engage the students to do more innovations and encourage production of their ideas.  He informed the delegation that during Covid-19, the government of Uganda provided funds to Makerere University, which were used to equip laboratories and do more research and produce a vaccine. The University also operationalized the online learning by use of technology to minimize the effects of the pandemic.  

The Vice Chancellor disclosed that there is an urgent need to re-tool the teachers in the Vocational institutes to upgrade their practical skills with the trends on the market. Therefore, the training and scholarship by Netherlands for vocational teachers to upgrade skills with latest technologies in universities would be appropriate.

Discussion with the College of Education and External Studies

The Deputy Principal, Dr. Ronald Bisaso received and welcomed the delegation. He represented the College Principal, Prof. Anthony Mugagga.  The Deputy Principal highlighted that regarding the education system in Uganda, some areas have changed and others improved.  He noted that many graduates lack the required skills for the job market. It would therefore be better if Makerere University also benefits from vocational studies and practice. Dr. Bisaso pointed out that the Department of Science, Technology and Vocation Education at the College of Education and External Studies offers a course on vocational studies and they expect to produce 1,500 graduates by 2025. The level of the vocational course offered is gauged by UBTEB (Uganda Business and Technical Examinations Board) that administers examinations and awards National Diplomas.  The investment in the education sector by government is quite minimal with just 11.5% (Higher education getting 6.4% and TVET getting 5.1%). Capacity of the sector needs to be enhanced through training. Professors must acquire entrepreneurial skills before they occupy leadership positions instead of doing so when they are already in the positions. At the College of Education and External Studies (CEES) with a population of 4,000 students, there only 30 doctoral students. CEES partners with the Ministry of Education and Sports through projects such as the Early childhood and development projects. Individual staff are seconded to projects to train and even share experiences.

The Deputy Principal called upon the Netherlands to support knowledge and capacity building of early career academics and partnering with the TVET ecosystem. This includes interventions, trainings and exchanges at various levels and cooperation with different stakeholders such as the government, the private sector, civil society and the Vocational institutes. He advocated for strengthening of existing vocational institutes, establishment of vocational institutes were they do not exist and development of research infrastructure and adoption of TVET across the education system.

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