Natural Sciences
Banana’s Large Plant Body Size an Advantage to Disease Control – Mak Scientists
Unlike for plants, in animals especially humans, body mass index (BMI, which is a person’s weight divided by the square of height) is a measure of physical health and pre-disposure to conditions like obesity. BMI does not make sense in plant health because of differences between plant and animal physiological systems.
However, large body size in plants may have some advantages. Apart from controlling a larger proportion of available resources and space within crowded vegetation, what other advantage does a large plant body size offer to an individual plant?
The banana’s plant body architecture
From the botanical point-of-view, the banana plant is a gigantic herb. A plant that is a herb or “herbaceous” is unable to undergo “secondary growth” and cannot form wood during its vegetative development.
The banana plant springs from an underground “true stem”, also called the “corm” or “rhizome”, to form a false stem, also called a “pseudostem” of 2-7 m height. The pseudostem is composed of the basal portions of leaf sheaths and is crowned with a rosette of 10 to 20 oblong to elliptic leaves that sometimes attain a length of 2-4 m and a breadth of 70 cm.
In mature banana plants, true stem emerges at the top of the pseudostem and bends downward to become a bunch of 10 to 300 individual fruits, or fingers, grouped in clusters, or hands, of 3 to 22. The edible part of the bunch is the female. In contrast, the inedible distal part, including the purple-colored cone-shaped end (locally known in some Ugandan dialects as “omukanaana” or “empumumpu”) constitutes the male part of the bunch.
How is the giant banana size an advantage in disease control?
A new study, titled “Xanthomonas campestris pv. musacearum Bacterial Infection Induces Organ-Specific Callose and Hydrogen Peroxide Production in Banana” and led by a team of scientists at the Department of Plant Sciences, Microbiology and Biotechnology at Makerere University in collaboration with the University of California, Davis, USA, shows how the giant banana body size can be used to control banana Xanthomonas wilt (BXW) disease.
According to Prof. Arthur Tugume, the lead scientist of this study and expert in plant pathology, when plants get infected, they respond instantly by implementing different strategies that limit the multiplication and/or mobility of the disease agents (pathogens). “For example, plants rapidly produce reactive oxygen species (ROS) such as hydrogen peroxide, superoxide ions, and hydroxyl ions. These ROS act as rapid messengers in the plant tissues to activate additional responses spreading over the entire plant body. This helps the plant’s distant tissues or organs to be aware and prepare advance defenses against the intruding pathogens”.
Prof. Dinesh-Kumar the project’s research collaborator based at the University of California-Davis, USA and expert in plant biology explains that “disease is a form of stress in plants and plants cannot perform well their biological functions when they are sick since they have to spend a lot of energy fighting against the disease. This is why disease control is important to enable plants grow well and yield high.”
The research indicates that ROS set in motion additional processes to ensure limited impact of disease and pathogens on the plant. For example, Hydrogen peroxide (H2O2), which is a ROS, has direct bactericidal, fungicidal or other anti-effects on the pathogens. Also, following H2O2 production, a unique plant carbohydrate, named “callose” starts to accumulate in large quantities within plant cells as a means of fortifying plant tissues. Callose differs from the other usual plant carbohydrates such as starch or cellulose because of the way its structures are formed.
Increased production of callose acts as a roadblock to any pathogen e.g., bacteria by limiting bacterial movement that would otherwise allow ease of attack on other tissues or cells at distant locations in the plant. “Although these plant defense responses are rapid, plant organs that are distant from the site of pathogen attack can be instrumental and block progression of bacteria or other pathogens by depositing callose in advance at strategic points” Prof. Tugume explains.
However, Prof. Tugume notes that callose participates also in many other normal developmental processes of plants, and for that reason, there is always some “housekeeping” callose in the plant tissues even without pathogenic infection. “This means that one must be able to accurately and quantitatively distinguish between ‘stress-induced’ and normal ‘housekeeping’ callose”, he adds.
How was the study done?
In this study, the researchers used young (2.5-months old) banana plantlets that had been generated from tissue culture at Kawanda Agricultural Research Institute. They then infected the plantlets with a bacterium called Xanthomonas campestris pv. musacearum (Xcm). This bacterium is the causative agent of banana Xanthomonas Wilt (BXW), the most destructive disease of bananas in East and Central Africa (ECA).
The banana leaves, pseudostems, corms and roots were analyzed for callose and compared with the control plants that had been inoculated with water instead of bacteria. H2O2 production was monitored by “DAB staining”, and by “spectrophotometry” while the analysis of callose was done by two methods: staining and visualization of callose using florescence microscopy, and using “Sandwich Enzyme-Linked Immunosorbent Assay” methods.
What did the researchers discover and how can it be used in BXW disease control?
This study revealed that the underground corm tissues assemble the stiffest resistance against BXW by depositing the highest concentrations of callose, while the pseudostem produced the highest quantities of H2O2. This is interesting for three (3) main reasons:
- Firstly, Xcm bacteria often enter through the leaves in regular plantation husbandry; hence, the corm being distant from leaves gives it an anatomical advantage in promoting the ability of lateral plants to escape Xcm infection.
- Secondly, the corm is an organ of perennation supporting vegetative and perennial continuity of the crop across seasons; hence it is charged in ensuring a disease-free next generation by severely constraining “mother-child transmission” of Xcm bacteria.
- Thirdly, the control of BXW now becomes easy when farmers are observant to the first aerial disease symptoms because Xcm is strongly constrained by bottlenecks in the pseudostem and corm.
Therefore, at the onset of aerial symptoms, diseased peudostems should immediately be removed by aseptically cutting them off at the corm without interfering with symptomless lateral shoots, which allows continuous food production and disease control to go on simultaneously. This is facilitated by the large size of the banana plant because at the onset of leaf symptoms (2.5 to 5 meters away from the corm), the bacteria have not yet arrived at the base of the pseudostem where the diseased plant can be cut off from the corm. This gives chance to a farmer to eliminate the infected pseudostems early (in 1 to 7 days) since the appearance of leaf symptoms.
This research was part of the PhD studies for Mr. Abubakar S. Mustafa at Makerere University and University of California, Davis. According to Mr. Mustafa, these discoveries make the management of BXW in banana plantations easy as long as the farmers are observant and act fast by removing diseased plants aseptically.
This study has been published by the American Phytopathological Society (APS) in an open access journal, “PhytoFrontiers”, and is freely accessible on https://apsjournals.apsnet.org/doi/full/10.1094/PHYTOFR-11-21-0073-R.
This study was funded by the Bill and Melinda Gates Foundation through the National Agricultural Research Organization (NARO), Uganda. The project had partners including the International Institute of Tropical Agriculture (IITA), the Alliance for Bioversity International and International Centre for Tropical Agriculture (CIAT) and Centre for Agriculture and Bioscience International (CABI).
For more details, contact;
Prof. Arthur Tugume
Lead Scientist
College of Natural Sciences (CoNAS)
Makerere University
Email: arthur.tugume@mak.ac.ug
Tel: +256772514841
Mr. Abubakar S. Mustafa
Co-Author and PhD student on the study
Email: mustafa.abubakar.sadik@gmail.com
Tel: +256702813233
Hasifa Kabejja
Principal Communication Officer
College of Natural Sciences (CoNAS)
Makerere University
Email: pr.cns@mak.ac.ug
Tel: +256774904211
Natural Sciences
Mak 76th Graduation Ceremony: CoNAS Presents 16 PhDs & Best Performing Male Student in the Sciences
The College of Natural Sciences (CoNAS) presented 269 students for the award of degrees of Makerere University at the 76th graduation ceremony. Of these, 16 graduated with PhDs (4 female and 12 male), 83 with Masters (27 female and 56 male), and 170 with Bachelor’s degrees (45 female and 125 male). The College also presented the Best Performing male student in the sciences. Simon Mungudit graduated with a CGPA of 4.76 in the Bachelor of Science in Petroleum Geoscience and Production. The PhD graduates included; AfazalI Zabibu, a Lecturer at the College of Business and Management Sciences (CoBAMS); Akuyenze Paul, Kawuma Carol, Mutiso Duncan Chalo, Nabiyonga Lydia, and Osinde Cyprian (Plant Sciences, Microbiology and Biotechnology); Batte Hebert, Katende Ronald, and Namugera Frank (Mathematics), Chaciga Jimmy, Tusiime Swaleh, and Komaketch Ignatius (Physics), Kinyua Mbuci, Sekandi Peter, and Shehu Zaccheus (Chemistry), and Nakiyende Herbert (Zoology, Entomology, and Fisheries Sciences).
On the second day of the four-day graduation ceremony, which began on 24th February 2026, three colleges presented students for the award of degrees and diplomas of Makerere University. These included CoNAS, the College of Health Sciences (CHS), and the College of Veterinary Medicine, Animal Resources and Biosecurity (COVAB).
Overall graduation Statistics
During the course of the 76th graduation ceremony, a total of 9,295 students will graduate with degrees and diplomas of Makerere University. Of these, 4,262 (46%) are female and 5,033 (54%) are male. A total of 213 students will graduate with PhDs, the highest number in the history of the University. 2,503 will graduate with Masters degrees; 6,343 with Bachelor’s degrees; 206 with postgraduate diplomas, and 30 with Diplomas.
Postgraduates constituted 31.4% of the total number of graduands.
Vice Chancellor’s Remarks
In his remarks, Vice Chancellor Prof. Barnabas Nawangwe commended CoNAS for its transformative research initiatives that are positively impacting communities, highlighting the NutriFishPlus Project, INNOECOFOOD Project, and a project to conserve medicinal plants in Eastern Uganda. Funded by the International Development Research Centre (IDRC) and Australian Centre for International Agricultural Research (ACIAR) under the Cultivate Africa’s Future Fund Phase II, the NutriFishPlus seeks to scale up fish processing technologies and empower fishing communities in Uganda.
The EU-funded INNOECOFOOD Project is transforming food production in Uganda. Through eco-innovative, climate-smart systems, the project aims to improve nutrition, protect the environment, and create jobs- especially for youth and women. Central to this effort are solar- and wind-powered ECOHUBs. These innovative centres use AI and IoT technology to support sustainable fish farming, monitor water quality, reduce losses, and increase yields.
Meanwhile, researchers from the Department of Plant Sciences, Microbiology, and Biotechnology, under the leadership of Dr. Patience Tugume, are spearheading efforts to conserve medicinal plants in Namutumba District. To date, 174 plant species have been documented, with ongoing assessments of threats from unsustainable harvesting. These plants play a vital role in treating malaria, diabetes, hypertension, and certain cancers.
Efforts to Transform Makerere into a Research-led University
In his address, the Vice Chancellor reaffirmed the University Management’s commitment to transforming Makerere into a research-led institution. He expressed gratitude to the Government of Uganda for securing a USD 162 million loan from the Korea Exim Bank to support critical infrastructure development at the University. The funding will facilitate the construction of new facilities for the School of Medicine, School of Dentistry, School of Engineering, the Science and Technology Centre, and the completion of the School of Computing and Information Sciences- an unprecedented development in the University’s history.
The Vice Chancellor further highlighted the pivotal role of the Makerere University Technology and Innovations Centre (MUTIC) in advancing incubation and commercialization. “The Centre offers mentorship, business development training, intellectual property support, and industry linkages to help transform research outputs into viable enterprises.” To date, eleven spin-off companies have been established by students and staff, while the University’s Innovation Pod (UniPod) has incubated more than 100 projects in the past year. The short-term target is to establish 50 spin-off companies annually, with plans to double that number in the long term.
Prof. Nawangwe acknowledged the government’s contribution through the Makerere University Research and Innovations Fund (MakRIF). Through this initiative, the government has supported over 1,400 high-impact research and innovation projects, and facilitated the training of more than 200 PhD students.
In partnership with the Science, Technology and Innovation Secretariat in the Office of the President, the University has also introduced awards to recognize outstanding researchers and innovators. These include the Research Excellence and Innovation Commercialization Awards. “The Innovation Commercialization Award underscores the institution’s commitment to translating research into practical products, services, and technologies that address real-world challenges and support national development,” the Vice Chancellor noted.
Vice Chancellor’s Message to the graduates and Appreciation to the Partners
The Vice Chancellor urged the graduates to uphold dignity, humility and diligence as they leave the gates of Makerere University, and begin the next chapter of their lives. “As you step out of the gates of Makerere, remain proud of your alma mater and serve as good ambassadors wherever you go. Use the knowledge acquired from one of the world’s leading universities to improve yourselves, your families, communities, country, and humanity at large.”
He equally appreciated the parents and guardians for supporting the students, and the development partners who have extended research funding to the university over the years. These include the Mastercard Foundation, the European Union, NORAD, Sida, IDRC, KOICA, DAAD, NIH, CDC, USAID, the Wellcome Trust, KfW, IUCEA, RUFORUM, ARUA, the PLUS Alliance, WUN, AAUN, AAP, the Madhvani Foundation, and the Government of Uganda through the Higher Education Students Financing Board. “We are equally grateful to the Embassy of China, the Chinese Chamber of Commerce, various Government Ministries, State House, private sector players, and many other partners whose generous support continues to advance our mission.”
Remarks by the Chancellor
Makerere University Chancellor, Hon. Dr. Crispus Kiyonga congratulated the graduands on their achievement and commended the staff, parents, and partners for their invaluable support. He expressed gratitude to the Government for its steadfast commitment to advancing research through the annual UGX30 billion allocation to the Makerere University Research and Innovations Fund.
He underscored the University’s contribution to Uganda’s Fourth National Development Plan (NDP IV) through advancements in science, technology, and innovation, and encouraged graduates to embrace entrepreneurship amidst an increasingly competitive job market. He further called for enhanced research funding, stronger private sector collaboration, and deeper community engagement to accelerate national development.
The Minister of Education and Sports
In her remarks, the Minister of Education and Sports, Hon. Janet Kataaha Museveni, represented by the Minister of State for Primary Education, Hon. Dr. Joyce Moriku Kaducu commended Makerere University for its pivotal role in driving national development. She highlighted the University’s contribution to knowledge generation and societal transformation. “The impressive number of PhD graduates this time round is testimony to the robust research output and academic excellence.” The Minister emphasized the importance of maintaining and enhancing academic standards, urging the University Administration to rigorously implement and uphold quality assurance measures that guarantee continued excellence of its programmes and global competitiveness of its graduates.
Implementing the Competence-based Education and Training
The Minister reiterated a key directive for all institutions of higher learning to prepare for the full implementation of Competence-Based Education and Training by July 2027. “This reform represents more than a policy shift. It signals a fundamental transformation in how graduates are prepared, with greater emphasis on practical skills, innovation, and problem-solving.” She urged the Vice Chancellor and his team to lead in curriculum reform, staff development, and infrastructure enhancement to ensure the successful rollout of competence-based education.
The Minister further urged the university management to ensure that all satellite campuses are fully accredited and that their programmes meet the same standards and relevance as those offered at the main campus.
Commencement Lecture by Dr. Maggie Kigozi
At the ceremony, distinguished alumnus, and Chair of the Makerere University Endowment Fund, Dr. Maggie Kigozi delivered a compelling commencement lecture, reflecting on her journey since earning her first degree in 1976. She shared three guiding messages for the new alumni as they embark on their professional paths.
Dr. Kigozi emphasized that power comes with responsibility, urging graduates entering the health and life sciences to combine their expertise with ethical leadership and financial literacy to create sustainable, compassionate practices. She highlighted the global opportunities that a Makerere degree offers, explaining how her education empowered her to make meaningful contributions across Uganda, Kenya, and Zambia.
She also encouraged the graduates to view failure as a stepping stone to excellence, noting that mistakes are not setbacks, but lessons. She underscored the importance of balance, reminding graduates to manage their personal and professional responsibilities realistically.
Research Excellence Awards
During the graduation ceremony, Makerere University recognized the top researchers and innovators from the 10 Colleges. At CoNAS, Mr. Omara Timothy from the Department of Chemistry scooped the Overall Top Research and Early Career Researcher Awards, whereas Dr Godwin Anywar from the Department of Plant Sciences, Microbiology, and Biotechnology won the Mid-Career Researcher Award.
More about Simon Mungudit, the best performing male science student – https://cns.mak.ac.ug/simon-mungudit-maks-best-male-science-student-this-year-rising-star-in-petroleum-geoscience/
Natural Sciences
BOKU University Charts New Collaboration Strategies with Mak’s Department of Zoology, Entomology & Fisheries Sciences
The Department of Zoology, Entomology and Fisheries Sciences at Makerere University on 11th February 2026 hosted a high-level delegation from the University of Natural Resources and Life Sciences, Vienna (BOKU), to discuss strategies for future collaboration in a partnership that has spanned five decades.
The meeting followed the BOKU team’s participation in the opening ceremony of the Promotion of Hybrid Renewable Energy Systems towards Electricity Access in Uganda project, an initiative designed to expand access to decentralized renewable energy solutions. The project, implemented under CEDAT, is funded by the Austrian Partnership Programme in Higher Education and Research for Development (APPEAR), a programme of the Austrian Development Cooperation administered by OeAD.
Discussions at the Department of Zoology, Entomology and Fisheries Sciences focused on deepening and broadening a historic collaboration that has shaped aquatic sciences, fisheries research, and environmental management in East Africa for half a century.
A Partnership Rooted in Limnology
The collaboration between Makerere University and BOKU traces its origins to 1975 with the establishment of the International Post-Graduate Course in Limnology (IPGL) by the late Heinz Löffler (1927-2006). The eight-month UNESCO-supported course, hosted in Lunz, Mondsee and Vienna, was designed to train scientists from developing countries in limnology – the study of inland waters.
In 1976, Dr. Gwahaba James Joshua of Makerere’s Department of Zoology, Entomology and Fisheries Sciences became the first beneficiary from the university. Two years later, Boniface Makanga attended the UNESCO course and went on to initiate a Marine Practical Course to the Indian Ocean in 1993. That programme has since grown into over 30 years of annual field activity in Mombasa, benefiting undergraduate and master’s students and strengthening hands-on marine science training at Makerere.
Over time, IPGL evolved into a vibrant International Joint Degree Master’s Programme in Limnology and Wetland Management. The programme has built capacity for 123 aquatic professionals – 56 female and 67 male, including 20 from Makerere University. Notably, 11 dedicated limnologists are currently based within the Department of Zoology, Entomology and Fisheries Sciences, a testament to sustained capacity development.
Expanding into Fisheries, Aquaculture and Regional Networks
The Mak-BOKU collaboration progressively expanded beyond limnology into fisheries, aquaculture, and broader freshwater ecosystem management.
Among the key initiatives was STRECAFISH (2015-2018), which focused on strengthening regional capacity in fisheries and aquaculture research and training to improve food security and livelihoods in Eastern Africa. The programme adopted a modular approach, trained 28 MSc students, supported five full PhDs and 11 MSc research projects, and facilitated 14 staff exchanges.
This momentum continued under COTRA (Collaborative Training in Fisheries and Aquaculture in East, Central and Southern Africa) from 2017 to 2023. COTRA enhanced staff and student mobility, supporting 24 master’s students (six credit-seeking and 18 degree-seeking), 12 doctoral candidates (four credit-seeking and eight degree-seeking), and 10 faculty and administrative staff. The initiative emphasized fit-for-purpose PhD training in aquaculture and fisheries to address food security challenges in Sub-Saharan Africa under the ACP-EU Cooperation Programme in Higher Education (EDULINK II), in partnership with institutions such as Lilongwe University of Agriculture and Natural Resources.
Ongoing collaborations include the Erasmus+ KA1 Learning Mobility of Individuals programme (2021-2027), which connects BOKU and Makerere with Egerton University, Kyambogo University, Addis Ababa University, Bahir Dar University and others. Additional joint initiatives include:
FreshNet: A higher education and research network for sustainable management of freshwater ecosystems in Eastern Africa, involving BOKU, Egerton University, Addis Ababa University, Bahir Dar University, EIAR and IHE Delft.
SWAQ-Uganda (2021-2026): Sustainable water quality management supporting Uganda’s development ambitions, implemented with IIASA, BOKU and Uganda’s Ministry of Water and Environment.
BIOGOLD (2024–2027): A project on biosorption for sustainable small-scale gold mining in Uganda, involving Kyambogo University, Makerere University, Gulu University, the University for Continuing Education and the University of Pretoria.
Collectively, these initiatives underscore a 50-year-old partnership that has built critical human resources at MSc and PhD levels in aquatic ecosystem management, conservation and utilization. The collaboration has also facilitated acquisition of essential research equipment, strengthened administrative capacity in project management, and nurtured a culture of knowledge sharing and innovation.
Meeting with the BOKU delegation
The BOKU delegation was led by the Rector, Prof. Eva Schulev-Steindl, a specialist in environmental law, and Vice Rector Assoc. Prof. Doris Damyanovic, an expert in landscape planning. They were accompanied by:
- Mag. Gerold Winkler, Coordinator of the Applied Limnology / Limnology and Wetland Master’s programme;
- Dr. Mathew Herrnegger, BOKU’s Africa Cooperation Country Coordinator and hydrology specialist; and
- Dr. Andreas Melcher, Africa-Uninet Coordinator from BOKU’s Institute for Development Research.
During the discussions with departmental staff, Dr. Akoll presented a detailed account of the partnership’s evolution and achievements, emphasizing its transformative impact on regional training and research capacity.
Charting the Next Chapter
As the partnership enters its sixth decade, both institutions are keen to expand into new frontiers. Key areas identified for future collaboration include:
- Enhanced staff and student exchanges;
- Joint research projects and co-publications;
- Introduction of short courses at the Makerere University Biological Field Station, Kibale beyond wildlife studies;
- Research on climate change and its effects on marine and freshwater resources;
- Advancement of basic biological research; and
- Strengthening fisheries research and innovation.
In his remarks, the Head of the Department of Zoology, Entomology and Fisheries Sciences, Dr. Godfrey Kawooya expressed gratitude for the enduring partnership, noting that it has significantly strengthened regional training capacity and supported the education of numerous Makerere staff and students. He added that the collaboration reinforces Makerere’s standing as one of the most collaborative universities globally and aligns closely with its research agenda.
Rector Schulev-Steindl reaffirmed BOKU’s commitment to advancing the longstanding relationship, describing it as a model of North-South academic cooperation built on mutual respect, shared knowledge and tangible impact.
Event pictorial: https://drive.google.com/drive/folders/1HkCCZf_TCPV0V7_ZR2PSVEw1zrEGhZeJ?usp=sharing
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